The List of Passalid Beetles from the Oriental Region (2)
LIST ONE-PART TWO
The List of the Passalid Beetles from the Oriental Region, including East Asia (part 2)
Subfamily Passalinae (continued)
* Leptaulax Kaup 1868 [Passalus dentatus Fabricius, by the subsequent designation, Zang 1905]
= Leptaulacides Zang 1905 [Type species: Passalus bicolor Fabricius, by the original designation]
= Trichostigmus Kaup 1871 [Type species: Leptaulax Thoreyi Kaup, monotypy]
* Leptaulax (Trichostigmus group = with hairy elytral margins)
* Leptaulax mindorensis (Boucher 1992) Philippines: Mindoro KI-Coll
* Leptaulax semperi (Boucher 1992) Philippines: Siargao near Mindanao --
* Leptaulax thoreyi Kaup 1868 Philippines: Luzon KI-Coll
? = Trichostigmus thoreyi, Boucher 1992 Philippines: Marinduque KI-Coll
? = Leptaulax sp. A (large)
* Leptaulax whiteheadi (Boucher 1992) Philippines: Luzon KI-Coll
@ Leptaulax sp. B (large) Philippines: Luzon KI-Coll
@ Leptaulax sp. C (large) Philippines: Negros KI-Coll
@ Leptaulax sp. D (small) Philippines: Marinduque KI-Coll
@ Leptaulax sp. E (small) Philippines: Negros KI-Coll
@ Leptaulax sp. F (small) Philippines: Panaon KI-Coll
@ Leptaulax sp. G (small) Philippines: Mindanao KI-Coll
* Leptaulax ursulus Schaufuss 1885 Sulawesi KI-Coll
* Leptaulax (Leptaulax group = with naked elytral margins)
* Leptaulax intermedius Iwase 1996 Philippines: Mindoro KI-Coll
* Leptaulax luzonicus Iwase 1996 Philippines: Luzon KI-Coll
* Leptaulax negrosensis Iwase 1996 Philippines: Negros KI-Coll
* Leptaulax sakaii Iwase 1996 Philippines: Negros KI-Coll
* Leptaulax arrowi Hincks, 1933
? = Leptaulax uenoi Iwase 1995 Philippines: Luzon KI-Coll
* Leptaulax immarginalis Iwase 1996 Sumatra KI-Coll
* Leptaulax immarginalis minangkabauensis Iwase 1998 Sumatra KI-Coll
* Leptaulax castilloae Kon, Johki & Araya, 2005 Malaya --
* Leptaulax masatakai Johki, Araya & Kon, 2003 Sumatra --
* Leptaulax punctatus Iwase 1995 Sumatra KI-Coll
* Leptaulax schoedli Iwase 1998 Sumatra --
* Leptaulax mindanaoensis Iwase 1996 Philippines: Mindanao KI-Coll
? Leptaulax anipunctatus (Zang 1905) Assam --
* Leptaulax cyclotaenius Kuwert 1891 Indochina, G. Sunda KI-Coll
= Leptaulax angustifrons Kuwert 1898
* ? Leptaulax himalayae Kuwert 1898 Indochina KI-Coll
(KI's L. himalayae from N Thailand: male genitalia resembles that of L. schillhammeri from G Sunda)
* Leptaulax kecil Iwase 1996 Borneo KI-Coll
* Leptaulax sabahensis Iwase 1996 Borneo KI-Coll
* Leptaulax schillhammeri Iwase 1998 Sumatra, Borneo, Malaya KI-Coll
(KI's some specimens -not all- of L. schillhammeri from Borneo: elytra almost shining, lacking worn-looking margin,
mesosternum with large punctures; they run down to L. cyclotaenius in my KEY on Leptaulax)
* Leptaulax dentifrons Iwase 1996 G. Sunda KI-Coll
* Leptaulax apicalis Iwase 1996 G. Sunda KI-Coll
* Leptaulax dentatus Fabricius 1792 (from Sumatra, ? complex) Taiwan, Indichina, G. Sunda KI-Coll
= Passalus quadridentatus Sturm 1826 (from *****)
= Leptaulax malitiosus Kuwert 1898 (from Java)
= Leptaulax submedius Kuwert 1898 (from Johore)
= Leptaulax indicus Kuwert 1891 (from Singapore; as var. of celebensis)
= Leptaulax insignis Kuwert 1898 (from Singapore; as ab. of indicus)
= Leptaulax mixtus Kuwert 1898 (from Borneo)
= Leptaulax celebensis var. bornensis Kuwert 1891 (from Borneo; as var. of celebensis)
= Leptaulax exterris Kuwert 1898 (from ? Brazil)
= Leptaulax saigonicus Kuwert 1891 (from Saigon)
= Leptaulax tonkinensis Zang 1905 (from Mt. Mauson)
= Leptaulax interponendus Kuwert 1898 (from China)
= Leptaulax darjelingi Kuwert 1891 (from Darjeeling)
= Leptaulax glabriventris Gravely 1914 (part, from Andaman, Java, Borneo)
* Leptaulax timoriensis Percheron 1841 (from Timor) L. Sunda (Timor) KI-Coll
L. Sunda (Sumbawa, Sumba, Flores) KI-Coll
= Leptaulax glabricollis Kuwert 1898 (from Timor)
? Leptaulax celebensis Kuwert 1891 (from Sulawesi) Philippines KI-Coll
Sulawesi KI-Coll
(male genitalia smaller than that of L. dentatus, lateral grooves of elytra narrower)
= Leptaulax separadus Kuwert 1891 (from the Philippines)
= Leptaulax geminatus Kuwert 1898 (from the Philippines)
= Leptaulax glabriventris Gravely 1914 (part, from the Philippines)
? Leptaulax ribbei Kuwert 1891 Sulawesi KI-Coll
(small, body not so flattened)
? Leptaulax sp. (close to celebensis, body smaller, strongly shining, elytra naked,
lamellae shorter, anterior lower tooth quadrangular) Sulawesi KI-Coll
* Leptaulax koreanus Nomura et al 1993 Korea --
* Leptaulax anna Zang 1905 L. Sunda KI-Coll
* Leptaulax beccarii Kuwert 1891 G. Sunda KI-Coll
* Leptaulax fujiokae Iwase 1995 Sumatra KI-Coll
* Leptaulax hidakai Johki, Araya & Kon 2003 Borneo --
* Leptaulax humerosus Kuwert 1891 G. Sunda KI-Coll
* Leptaulax laevis Iwase 1998 Sumatra --
* Leptaulax masayukii Iwase 1997 L. Sunda: Sumbawa KI-Coll
L. Sunda: Flores --
* Leptaulax arayai Johki & Kon 2003 G. Sunda (Borneo) KI-Coll
* Leptaulax elegans Iwase 1996 G. Sunda KI-Coll
* Leptaulax acutangularis Iwase 1998 G. Sunda KI-Coll
? Leptaulax cicatricosus Kuwert 1891 (from Seram(1891) or Sulawesi(1898) Sulawesi KI-Coll
* Leptaulax crockerensis Iwase 1998 G. Sunda (Borneo) --
* Leptaulax jaechi Iwase 1998 Sulawesi --
* Leptaulax jenisi Iwase 1998 G. Sunda (Borneo) KI-Coll
* Leptaulax labyrinthus Iwase 1998 Sulawesi KI-Coll
? Leptaulax macassariensis, Kuwert 1898 (nec Schaufuss) Sulawesi KI-Coll
* Leptaulax parvus Iwase 1995 G. Sunda (Sumatra) KI-Coll
* Leptaulax parvus balita Iwase 1998 G. Sunda (Borneo) KI-Coll
* Leptaulax strbai Iwase 1998 G. Sunda (Borneo) --
* Leptaulax trajae Iwase 1998 Sulawesi KI-Coll
@ Leptaulax sp. of Cameron Highlnds P. Malaysia KI-Coll
* Leptaulax matsumotoi Kon, Johki & Araya 2003 Laos KI-Coll
* Leptaulax matsumotoi ssp. kangfangensis Kon & Abe, 2009 Myanmar --
* Leptaulax riekoae Iwase, 1998 Thailand, Laos KI-Coll
* Leptaulax riekoae ssp. kachinensis Kon, Araya & Johki, 2005 Myanmar --
* Leptaulax roepstorfi Kuwert 1898 Andaman --
= Leptaulax planicollis Kuwert 1898
? = Leptaulax andamanarum (Zang 1905) Andaman KI-Coll
(L. roepstorfi of Reyes-Castillo & Castillo 1986 = L. andamanarum of KI-Coll)
* Leptaulax planus Illiger 1800 G. Sunda KI-Coll
* Leptaulax barbicauda (Zang 1905) (sensu Gravely, 1918) G. Sunda KI-Coll
? = Leptaulax anaulax (Zang 1905) (from Sumatra)
? = Leptaulax niae Kuwert 1898 (from Nias)
? = Leptaulax palawanicus (Zang 1905) (from Palawan)
? = Leptaulax sp. K Philippines: Luzon, Negros, Mindanao
= Leptaulax eschscholtzi, Kuwert, 1898 (nec Kaup)
(L. barbicauda is only recorded from Peninsular Malaysia in Gravely's key, but some forms are known in KI-Coll,
they are charaterized by the flattened body, the hairy shoulders and the mandibular teeth, the distal abdominal
sternite is variable, hairy or naked, sometimes shows the sexual dimorphism)
* Leptaulax eschscholtzi Kaup 1868 (from Nias)
* Leptaulax loebli Kon, Johki & Araya, 2003 Thailand, Myanmar KI-Coll
* Leptaulax sambawae Gravely 1918 L. Sunda (Sumbawa) KI-Coll
* Leptaulax bicolor Fabricius 1801, sensu Gravely 1914~1918 KI-Coll
(L. bicolor sensu Gravely includes many different kinds of species, they are charaterized as follows:
1, hypostomal process smooth and flat (probably),
2, parietal ridge extending to supraorbital ridge (probably)
3, frontal area naked (probably),
4, elytral shoulders naked, lacking hair tufts (probably),
5, Eyes not so small (probably)
6. integment of middle and hind tibiae never worn-looking (probably)
6, Not Leptaulax manis Iwase (probably, and this is recorded from Indochina, G. Sunda & Sulawesi now)
(Malay, Sumatra, Borneo popul.) G. Sunda KI-Coll
? = Leptaulax abdominibarbatus Kuwert 1898 (from Sumatra)
? = Leptaulax anibarbis Kuwert 1898 (from Borneo)
? = Leptaulax vicinus Percheron 1844 (from *****)
? = Leptaulax innocuus DEJEAN 1837 (from *****)
? = Leptaulax sumatrae Kuwert 1898 (from Sumatra)
? = Leptaulax geminus Kuwert 1898 (from Sinnep)
(Java popul)
? = Leptaulax sequens Kuwert 1898 (from Java)
@ Leptaulax sp. from W. Java Java W --
@ Leptaulax sp. from C. Java Java C KI-Coll
@ Leptaulax sp. from E. Java Java E KI-Coll
(Bali popul)
@ Leptaulax sp. (25 mm~) Bali KI-Coll
(Philippine popul: three L. bicolor-like species in KI-Coll) Philippines KI-Coll
? = Leptaulax differentispina Kuwert 1891 (probably valid)
? = Leptaulax separandus var. medius Kuwert 1891
? = Leptaulax medius var. divaricatus Kuwert 1898
(Sulawesi popul): some undescribed species in Sulawesi Sulawesi KI-Coll
? = Leptaulax consequens Kuwert
? = Leptaulax macassariensis Schaufuss 1885
(Indochina popul) Indochina~Myanmar KI-Coll
? = Leptaulacides fruhstorferi Zang 1905 (from Than Moi & Mt. Mauson)
? = Leptaulacides rugulosus Zang 1905 (from Mt. Mauson)
? = Leptaulax tenesserimensis Kuwert 1898 (from Tenasserim)
@ Leptaulax sp. B of Indochina Vietnam C-H KI-Coll
@ Leptaulax sp. C of Indochina Laos KI-Coll
(India popul.) India --
? = Leptaulax dindigalensis Kuwert 1898 (from Dindigal)
? = Leptaulax subsequens Kuwert 1891 (from E. India (1891), Calcutta, S. Sulawesi & Seram (1898))
? = Leptaulax incipiens Kuwert 1891 (from E. India (1891), Andaman & E. India (1898))
? = Leptaulax abdominisculptus var. calcuttae Kuwert 1891 (from Calcutta)
? = Leptaulax aurivillii Kuwert 1891 (from Assam)
? = Leptaulax evidens Kuwert 1898 (from Assam)
(Sri Lanka popul.) Sri Lanka --
? = Leptaulax nietneri Zang 1905 (from Sri Lanka)
* Leptaulax malaccae Kuwert 1891 G. Sunda KI-Coll
* Leptaulax manillae Kuwert 1891 Philippines KI-Coll
* Leptaulax manis Iwase 1996 Indochina: Thailand KI-Coll
G. Sunda KI-Coll
Philippines: Basilan
Sulawesi KI-Coll
* Leptaulax similis Iwase 1996 Borneo KI-Coll
* Leptaulax formosanus Doesburg 1992 Taiwan KI-Coll
* Leptaulax pacholatkoi Iwase 1998 Vietnam C-H KI-Coll
* Leptaulax sarawakus Iwase 1996 Borneo KI-Coll
@ Leptaulax sp. A of Indochina (related to L. pacholatkoi) Laos KI-Coll
@ Leptaulax sp. D of Indochina Vietnam KI-Coll
* Leptaulax datar Iwase 1996 Malaya KI-Coll
* Leptaulax karoensis Iwase 1996 Sumatra N --
* Leptaulax iwasei Johki, Araya & Kon, 2007 Sumatra W --
* Leptaulax circulus Iwase 1996 Borneo KI-Coll
* Macrolinus Kaup 1868 [Type species: Passalus latipennis by the subsequent designation, Gravely 1918]
* Macrolinus crenatipennis Kuwert 1898 Sri Lanka --
* Macrolinus obesus Gravely 1918 Sri Lanka --
* Macrolinus rotundifrons Kaup 1871 Sri Lanka --
= Macrolinus singhalensis Kuwert 1898
* Macrolinus waterhousei Kaup 1871 Sri Lanka --
* Macrolinus andamanensis Stoliczka 1873 Andaman KI-Coll
* Macrolinus nicobaricus Gravely 1914 Nicobar --
* Macrolinus sikkimensis STOLICZKA 1873 E. India KI-Coll
Indochina KI-Coll
= Macrolinus tavoyanus Gravely 1914 (from Tavoy & Dawna)
* Macrolinus batesi Kuwert 1898 Sumatra KI-Coll
* Macrolinus depressus Gravely 1918 Java --
* Macrolinus mineti Boucher 1996 Sumatra ? KI-Coll
* Macrolinus cartereti Boucher 1996 Malaya KI-Coll
* Macrolinus puncticollis HINCKS 1938 Java --
* Macrolinus latipennis Percheron 1841 (from Java; ! complex) G. Sunda KI-Coll
Philippines KI-Coll
= Macrolinus marginepunctatus DEJEAN 1837
= Macrolinus weberi STOLICZKA 1873 (rec. from Johore;nec Kaup)
= Macrolinus orbatus Kuwert 1898 (from Java)
= Macrolinus avanus Kuwert 1898 (from Java)
= Macrolinus dissimilis Kuwert 1898 (from Nias)
= Macrolinus latipennis ab. fraternus Kuwert 1898 (from Nias, the Philippines & Moluku)
= Macrolinus parallelipennis Kuwert 1898 (from Java)
* Macrolinus weberi Kaup 1868 Philippines KI-Coll
* Macrolinus duivenbodei Kaup 1868 Sulawesi --
Tahulandang KI-Coll
* Macrolinus salibabuensis Boucher 1996 Sangihe --
* Macrolinus punctipectus HINCKS 1838 Sulawesi --
* Macrolinus sulciperfectus Kuwert 1891 (? complex) Sulawesi KI-Coll
* Macrolinus urus HELLER 1898 Sulawesi --
* Ophrygonius Zang 1904 [Type species: Ophrygonius quadrifer Zang, monotypy]
* Ophrygonius inaequalis (Burmeister 1847) G. Sunda KI-Coll
= oroleius SMITH 1852 (from ? Korea)
= sinkepicus Kuwert 1898 (from Sinkep Is.)
= Ophrygonius quadrifer Zang 1904
* Ophrygonius planus Iwase 1995 G. Sunda --
* Aceraius Kaup 1868 [Type species: Passalus grandis BURMEISTER, by the subsequent designation, Gravely 1918]
= Basilianus Kaup 1871 [Type species: probably not designated, neelgherriensis is the first example in Kaup, 1871]
= Heterochilus Kuwert 1891 [Type species: Heterochilus crinitus Kuwert, monotypy]
= Episphenus Kaup 1871 [Type species: Episphenus Moorei Kaup, monotypy]
= Laches Kaup, 1871 [Type species: Aceraius Comptoni Kaup, ? designated by HINCKS & DIBB]
= Chilomazus Zang [n.n. for Laches]
= Cacoius Boucher 1993 [Type species: Cacoius jacquesi Boucher, original monotypy]
* Aceraius (group Chilomazus)
* Aceraius comptoni Kaup 1868
= Chilomazus comptoni (Kaup 1868)
= Episphenus comptoni (Kaup) Sri Lanka --
= Laches fruhstorferi Kuwert 1891
= Laches puella Kuwert 1891
= Laches parallelogrammifrons Kuwert 1898
= Laches socius Kuwert 1898
* Aceraius flachi (Kuwert 1891)
= Chilomazus flachi (Kuwert 1891)
= Episphenus flachei (Kuwert, 1891) Sri Lanka --
* Aceraius sp. (pronotum with several hairs in scars) Sri Lanka -- (Wien N-H Mus)
* Aceraius (group Episphenus)
* Aceraius moorei (Kaup 1871) Sri Lanka KI-Coll (Hong Kong)
= Episphenus moorei Kaup, 1871
= Episphenus pearsoni Gravely 1914
* Aceraius annamensis (Gravely 1919) Indochina --
= Episphenus annamensis Gravely, 1919
* Aceraius (group Basilianus), incl. Cacoius Boucher, 1993 [If the group is divided from Aceraius group, Basilianus should be
used for the group name]
[HAIRlESS ELYTRAL GROUP]
* Aceraius boxeri (Boucher 1993) S. India KI-Coll
= Ophrygonius boxer Boucher, 1993
* Aceraius indicus (Stoliczka 1873) S. India --
= Basilianus indicus Stoliczka 1873
= Aceraius cantori Kaup 1868 (pars, from Malabar; nec Percheron)
= Aceraius neelgheriensis Kaup (pars, from Malabar)
= Basilianus certus Kuwert 1891 (from E. India)
= Basilianus stoliczkae Kuwert 1891 (from E. India)
* Aceraius neelgherriensis (Percheron 1841) S. India KI-Coll
= Basilianus neelgherriensis (Percheron, 1841)
= Ophrygonius neelgherriensis (Percheron, 1841)
= Aceraius infantilis Kaup 1868 (from Vanicoro)
= Aceraius puerilis Kaup 1868 (from Aru)
= Basilianus nilgheriensis Kuwert 1891 (rec. from Malabar)
= Analaches filius Kuwert 1898 (from Nilgiri)
= Basilianus inaequalis, Kuwert 1898 (rec. from Malacca & Dindigal; nec BURMEISTER)
= Basilianus certus Kuwert, 1898 (from Malabar; see indicus)
= Basilianus binominis Zang 1905 (from Dindigul)
* Aceraius cantori (Percheron 1844) India NE KI-Coll
= Ophrygonius cantori (Percheron, 1844)
= Basilianus interrogationis Kuwert 1891
? = Ophrygonius cantori var. dunsiriensis Gravely, 1914 Assam KI-Coll
* Aceraius chinensis (Endroedi 1955, nec Kuwert) China KI-Coll
= Ophrygonius cantori chinensis Endoedi, 1955
= Ophrygonius chinensis Endrodi, 1955 (originally described as O. cantori chinensis, the combination with Aceraius
is the secondary homonym, with Basilianus is available)
* Aceraius convexifrons (Zang 1904) Assam --
[should be divided into some local forms]
Indochina (N.Vietnam) KI-Coll
Indochina (N.Thailand)KI-Coll
S.China KI-Coll
* Aceraius uedai (Kon & Johki 1991) Borneo KI-Coll
= Ophrygonius uedai Kon & Johki 1999
* Aceraius emas Iwase 1998 Borneo --
= Ophrygonius emas (Iwase 1998)
* Aceraius rajanus (Doesburg 1941) ? Malaya --
= Ophrygonius rajanus Doesburg 1941
* Aceraius inopinus (Boucher 1997) S. Thailand --
= Ophrygonius inopinus Boucher 1997
* Aceraius lacroixi (Boucher 1993) Sumatra --
= Ophrygonius lacroixi Boucher 1993
* Aceraius longiruga (Boucher 1993) Sumatra KI-Coll
= Ophrygonius longiruga Boucher 1993
* Aceraius bleuzeni (Boucher 1993) Malaya --
= Ophrygonius longiruga Boucher 1993
* Aceraius birmanicus (Gravely 1914) Indochina, Myamar KI-Coll
= Ophrygonius birmanicus Gravely 1914
* Aceraius ferreri (Boucher 1993) Malaya KI-Coll
= Ophrygonius ferreri Boucher 1993
* Aceraius koni (Boucher 1995) Borneo KI-Coll
= Ophrygonius koni Boucher 1995
= Aceraius symmetricus Iwase 1995
* Aceraius singapurae (Gravely 1914) G. Sunda KI-Coll
= Ophrygonius singapurae Gravely, 1914
[HAIRY ELYTRAL GROUP]
* Aceraius minor Gravely 1914 Malaya KI-Coll
= Ophrygonius minor (Gravely 1914)
= Aceraius parvulus Hincks, 1938
* Aceraius tuberculosus (Boucher & Kon, 1999) Malaya --
= Ophrygonius tuberculosus Boucher & Kon, 1999
* Aceraius tuberculus (Boucher 1995) Malaya --
= Ophrygonius tuberculus Boucher 1995
* Aceraius aequalis (Gravely 1918) N. Vietnam KI-Coll
= Ophrygonius aequalis Gravely 1918
* Aceraius javensis (Gravely 1918) Java --
= Ophrygonius javensis Gravely, 1918
* Aceraius megatarsi (Boucher & Kon, 1999) Sumatra --
= Ophrygonius megatarsi Boucher & Kon, 1999
* Aceraius pseudaequidens (Boucher 1995) Borneo --
= Ophrygonius pseudaequidens Boucher 1995
* Aceraius sternibarbis (Boucher 1995) Borneo --
= Ophrygonius sternibarbis Boucher 1995
* Aceraius griveaui (Boucher 1993) N. Vietnam --
= Ophrygonius griveaui Boucher 1993
* Aceraius tanae (Boucher 1993) C. Vietnam --
= Ophrygonius tanae Boucher 1993
* Aceraius aequidens Gravely 1914 Borneo KI-Coll
= Ophrygonius aequidens (Gravely 1914)
* Aceraius hirtimarginalis Iwase 1993 Borneo KI-Coll
= Ophrygonius hirtimarginalis (Iwase 1993)
* Aceraius jacquesi (Boucher 1993) Borneo KI-Coll
= Cacoius jacquesi Boucher 1993
* Aceraius (group Aceraius)
* Aceraius alutaceosternus Kuwert 1898 G. Sunda KI-Coll
* Aceraius helferi Kuwert 1891 G. Sunda, Indochina KI-Coll
Korea --
India NE --
= Aceraius pilifer, Kuwert 1898 (nec Percheron)
= Aceraius tavoyanus Gravely 1914
= Aceraius emarginatus, Stoliczka (nec Weber; nec Percheron)
? = Aceraius assamensis Kuwert 1898 (from Assam)
* Aceraius himalayensis Gravely 1914 Indochina KI-Coll
* Aceraius borneanus Kaup 1871 G. Sunda KI-Coll
= Aceraius Kaupi Kirsch in Kaup 1871
= Aceraius Percheroni Kaup 1871
= Aceraius incidens Kirsch 1875
= Aceraius nanus Kuwert 1898
* Aceraius lamellatus Gravely 1918 (complex) G. Sunda KI-Coll
Philippines KI-Coll
Sulawesi KI-Coll
* Aceraius laniger Zang 1905 Borneo KI-Coll
* Aceraius kikutai KON et al 1995 Borneo KI-Coll
* Aceraius Boucheri KON et al 1993 Borneo KI-Coll
= Aceraius moeschleri, KON et al 1992 (nec Kuwert)
* Aceraius katsurae Iwase 1995 Borneo KI-Coll
* Aceraius jenisi Iwase 1998 Borneo --
* Aceraius riekoae Iwase 1998 Borneo KI-Coll
* Aceraius hidakai KON et al 1993 Borneo KI-Coll
* Aceraius laevimargo Zang 1905 Borneo KI-Coll
* Aceraius wallacei (Kuwert 1898) G. Sunda KI-Coll
= Heterochilus crinitus Kuwert 1891 (from Nias)
= Heterochilus oculitessellatus Kuwert 1898 (from Nias)
= Aceraius perakensis Kuwert 1898 (! nec Gravely)
* Aceraius pilifer Percheron 1835 Java KI-Coll
= Aceraius emarginatus, Kuwert 1898 (nec Weber, nec Percheron)
= Aceraius reticulaticollis Kuwert 1898
? = Aceraius nikobaricus (Redtenbacher 1867)
* Aceraius tobae Iwase 1994 [should divided into some species] Sumatra KI-Coll
* Aceraius kazuhisai Kon, Johki & Araya, 2003 Sumatra KI-Coll
= Aceraius ashidai, Iwase 1994 (part; nec KON et al)
* Aceraius masatakai Kon, Johki & Araya, 2003 Sumatra KI-Coll
* Aceraius fujiokae Iwase 1994 Sumatra KI-Coll
* Aceraius parvulus HINCKS 1938 Malaya --
* Aceraius ashidai KON et al 1992 Malaya KI-Coll
* Aceraius hikidai KON et al 1993 Borneo KI-Coll
? = Aceraius borneanus, Kuwert 1898 (nec Kaup)
* Aceraius sabanus KON et al 1995 Borneo KI-Coll
* Aceraius alpinus KON et al 1995 Borneo --
* Aceraius hassani Kon, Araya & Maryati (Mohamed), 2006 Borneo KI-Coll
* Aceraius tricornis Zang 1903 Borneo KI-Coll
* Aceraius kinabalensis KON & JOHKI 1989 Borneo KI-Coll
* Aceraius Kuwerti Zang 1903 Borneo KI-Coll
* Aceraius laevicollis (Illiger 1800 ) G. Sunda KI-Coll
= Passalus emarginatus Weber 1801
= Aceraius meyeri Kuwert 1891 (from Singapore)
= Aceraius pontifex Kuwert 1898 (from Malacca)
* Aceraius luzonicus Kuwert 1898 Philippines KI-Coll
* Aceraius sipolae Iwase 1998 Sipola near Sumatra KI-Coll
* Aceraius moescheri Kuwert 1891 G. Sunda KI-Coll
* Aceraius illegalis Kuwert 1891 G. Sunda KI-Coll
* Aceraius grandis BURMEISTER 1847 (from Java) G. Sunda KI-Coll
= [Aceraius grandis auct. (complex); this relative from Taiwan is unnamed]
= Passalus emarginatus, Percheron 1835 (rec. from Java; nec Weber)
= Aceraius addendus Kuwert 1891 (from Java)
= Aceraius rectidens Kuwert 1891 (from the Philippines, but Malacca & Sumatra in 1898)
= Aceraius grandis ab. minutifrons Kuwert 1898
= Aceraius prosternisulcatus Kuwert 1898 (from Moluku)
= Aceraius magnus Kuwert 1898 (from Sumatra)
= Aceraius molossus Kuwert 1898 (from Java)
? Aceraius palawanus Zang 1905 (from S. Palawan) Philippines KI-Coll
? Aceraius hirsutus Kuwert 1891 (from Sikkim) Indochina KI-Coll
? = Aceraius ceylonicus Kuwert 1891 (from Sri Lanka)
? Aceraius chinensis Kuwert 1898 (from S. China; nec Endroedi) S. China KI-Coll
* Aceraius oculidens Zang 1905 G. Sunda KI-Coll
* Aceraius (NOT grouping) sawaii Kon, Araya & Maryati (Mohamed), 2006 Borneo --
* : valid species
= : synonymic relation
@ (genus) sp. : probably unnamed or not described species
? = (genus) sp. : determined by KI, specific name uncertain (probability of 'stat. rev.')
? (genus) sp. : determined by KI, probability of 'stat. rev.'
2009/09/25
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